The house sparrow ( Passer domesticus) is a bird of the sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, and males have brighter black, white, and brown markings. One of about 25 species in the genus Passer, the house sparrow is native to most of Europe, the Mediterranean Basin, and a large part of Asia. Its intentional or accidental introductions to many regions, including parts of Australasia, Africa, and the Americas, make it the most widely distributed wild bird.
The house sparrow is strongly associated with human habitation, and can live in urban or rural settings. Though found in widely varied habitats and climates, it typically avoids extensive , , polar regions, and hot, dry far away from human development. For sustenance, the house sparrow routinely feeds at home and public bird feeding stations, but naturally feeds on the seeds of grains, flowering plants and weeds. However, it is an opportunistic, omnivorous eater, and commonly catches invertebrates such as insects and their larvae, , and many other natural foods.
Because of its numbers, ubiquity, and association with human settlements, the house sparrow is culturally prominent. It is extensively, and usually unsuccessfully, persecuted as an agricultural pest. It has also often been kept as a pet, as well as being a food item and a symbol of lust, sexual potency, commonness, and vulgarity. Though it is widespread and abundant, its numbers have declined in some areas. The bird's conservation status is listed as least concern on the IUCN Red List.
In mass, the house sparrow ranges from . Females usually are slightly smaller than males. The median mass on the European continent for both sexes is about , and in more southerly subspecies is around . Younger birds are smaller, males are larger during the winter, and females are larger during the breeding season. Birds at higher latitudes, colder climates, and sometimes higher altitudes are larger (under Bergmann's rule), both between and within subspecies.
The male is duller in fresh nonbreeding plumage, with whitish tips on many feathers. Wear and preening expose many of the bright brown and black markings, including most of the black throat and chest patch, called the "bib" or "badge". The badge is variable in width and general size, and may signal social status or fitness. This hypothesis has led to a "veritable 'cottage industry of studies, which have only conclusively shown that patches increase in size with age. The male's bill is dark grey, but black in the breeding season.
The female has no black markings or grey crown. Its upperparts and head are brown with darker streaks around the mantle and a distinct pale supercilium. Its underparts are pale grey-brown. The female's bill is brownish-grey and becomes darker in breeding plumage approaching the black of the male's bill.
Juveniles are similar to the adult female, but deeper brown below and paler above, with paler and less defined supercilia. Juveniles have broader buff feather edges, and tend to have looser, scruffier plumage, like moulting adults. Juvenile males tend to have darker throats and white postoculars like adult males, while juvenile females tend to have white throats. However, juveniles cannot be reliably sexed by plumage: some juvenile males lack any markings of the adult male, and some juvenile females have male features. The bills of young birds are light yellow to straw, paler than the female's bill. Immature males have paler versions of the adult male's markings, which can be very indistinct in fresh plumage. By their first breeding season, young birds generally are indistinguishable from other adults, though they may still be paler during their first year.
Aggressive males give a trilled version of their call, transcribed as "chur- chur-r-r-it-it-it-it". This call is also used by females in the breeding season, to establish dominance over males while displacing them to feed young or incubate eggs. House sparrows give a nasal alarm call, the basic sound of which is transcribed as quer, and a shrill chree call in great distress. Another vocalisation is the "appeasement call", a soft quee given to inhibit aggression, usually given between birds of a mated pair. These vocalisations are not unique to the house sparrow, but are shared, with small variations, by all sparrows.
The bird's scientific name and its usual English name have the same meaning. The Latin word passer, like the English word "sparrow", is a term for small active birds, coming from a root word referring to speed. The Latin word domesticus means "belonging to the house", like the common name a reference to its association with humans. The house sparrow is also called by a number of alternative English names, including English sparrow, chiefly in North America; and Indian sparrow or Indian house sparrow, for the birds of the Indian subcontinent and Central Asia. Dialectal names include sparr, sparrer, spadger, spadgick, and philip, mainly in southern England; spug and spuggy, mainly in northern England; spur and sprig, mainly in Scotland; and spatzie or spotsie, from the German Spatz, in North America.
The taxonomy of the house sparrow and its Mediterranean relatives is complicated. The common type of "willow sparrow" is the Spanish sparrow, which resembles the house sparrow in many respects. It frequently prefers wetter habitats than the house sparrow, and it is often colonial and nomadic. In most of the Mediterranean, one or both species occur, with some degree of hybridisation. In North Africa, the two species hybridise extensively, forming highly variable mixed populations with a full range of characters from pure house sparrows to pure Spanish sparrows.
In most of Italy, the breeding species is the Italian sparrow, which has an appearance intermediate between those of the house and Spanish sparrows. Its specific status and origin are the subject of much debate, but it may be a case of long-ago hybrid speciation. In the Alps, the Italian sparrow over a narrow roughly strip with the house sparrow, and some house sparrows migrate into the Italian sparrow's range in winter. On the Mediterranean islands of Malta, Gozo, Crete, Rhodes, and Karpathos, other apparently intermediate birds are of unknown status.
Migratory birds of the subspecies P. d. bactrianus in the P. d. indicus group were recorded overlapping with P. d. domesticus birds without hybridising in the 1970s, so the Soviet scientists Edward I. Gavrilov and M. N. Korelov proposed the separation of the P. d. indicus group as a separate species. However, P. d. indicus group and P. d. domesticus group birds intergrade in a large part of Iran, so this split is rarely recognised.
In North America, house sparrow populations are more differentiated than those in Europe. This variation follows predictable patterns, with birds at higher latitudes being larger and darker and those in arid areas being smaller and paler. However, how much this is caused by evolution or by environment is not clear. Similar observations have been made in New Zealand and in South Africa. The introduced house sparrow populations may be distinct enough to merit subspecies status, especially in North America and southern Africa, and American ornithologist Harry Church Oberholser even gave the subspecies name P. d. plecticus to the paler birds of western North America.
The first of many successful introductions to North America occurred when birds from England were released in New York City, in 1852, intended to control the ravages of the Ellida caniplaga. In North America, the house sparrow now occurs from the Northwest Territories of Canada to southern Panama, and it is one of the most abundant birds of the continent. The house sparrow was first introduced to Australia in 1863 at Melbourne and is common throughout the eastern part of the continent as far north as Cape York, but has been prevented from establishing itself in Western Australia, where every house sparrow found in the state is killed. House sparrows were introduced in New Zealand in 1859, and from there reached many of the Pacific islands, including Hawaii.
In southern Africa, birds of both the European subspecies ( P. d. domesticus) and the Indian subspecies ( P. d. indicus) were introduced around 1900. Birds of P. d. domesticus ancestry are confined to a few towns, while P. d. indicus birds have spread rapidly, reaching Tanzania in the 1980s. Despite this rapid spread, native relatives such as the Cape sparrow also occur and thrive in urban habitats. In South America, it was first introduced near Buenos Aires around 1870, and quickly became common in most of the southern part of the continent. It now occurs almost continuously from Tierra del Fuego to the fringes of the Amazon basin, with isolated populations as far north as coastal Venezuela.
It tolerates a variety of climates, but prefers drier conditions, especially in moist tropical climates. It has several adaptations to dry areas, including a high salt tolerance and an ability to survive without water by ingesting berry. In most of eastern Asia, the house sparrow is entirely absent, replaced by the Eurasian tree sparrow. Where these two species overlap, the house sparrow is usually more common than the Eurasian tree sparrow, but one species may replace the other in a manner that ornithologist Maud Doria Haviland described as "random, or even capricious". In most of its range, the house sparrow is extremely common, despite some declines, but in marginal habitats such as rainforest or mountain ranges, its distribution can be spotty.
Several studies of the house sparrow in temperate agricultural areas have found the proportion of seeds in its diet to be about 90%. It will eat almost any seeds, but where it has a choice, it prefers cereals: , wheat or maize. Rural birds tend to eat more waste seed from animal dung and seed from fields while urban birds tend to eat more commercial bird seed and weed seed. In urban areas, the house sparrow also feeds largely on food provided directly or indirectly by humans, such as bread, though it prefers raw seeds. The house sparrow also eats some plant matter besides seeds, including , berries, and fruits such as grapes and cherries. In temperate areas, the house sparrow has an unusual habit of tearing flowers, especially yellow ones, in the spring.
Animals form another important part of the house sparrow's diet, chiefly , of which , , flies, and are especially important. Various noninsect are eaten, as are and where available, , and even such as and . Young house sparrows are fed mostly on insects until about 15 days after hatching. They are also given small quantities of seeds, spiders, and grit. In most places, and crickets are the most abundant foods of nestlings. Hemiptera, , sawfly, and beetles are also important, but house sparrows take advantage of whatever foods are abundant to feed their young. House sparrows have been observed Kleptoparasitism prey from other birds, including .
The gut microbiota of house sparrows differs between chicks and adults, with Pseudomonadota (formerly Proteobacteria) decreasing in chicks when they get to around 9 days old, whilst the relative abundance of Bacillota increase.
Males take up nesting sites before the breeding season, by frequently calling beside them. Unmated males start nest construction and call particularly frequently to attract females. When a female approaches a male during this period, the male displays by moving up and down while drooping and shivering his wings, pushing up his head, raising and spreading his tail, and showing his bib. Males may try to mate with females while calling or displaying. In response, a female will adopt a threatening posture and attack a male before flying away, pursued by the male. The male displays in front of her, attracting other males, which also pursue and display to the female. This group display usually does not immediately result in copulations. Other males usually do not copulate with the female. Copulation is typically initiated by the female giving a soft dee-dee-dee call to the male. Birds of a pair copulate frequently until the female is laying eggs, and the male mounts the female repeatedly each time a pair mates.
The house sparrow is monogamous, and typically mates for life, but birds from pairs often engage in extra-pair copulations, so about 15% of house sparrow fledglings are unrelated to their mother's mate. Males guard their mates carefully to avoid being cuckolded, and most extra-pair copulation occurs away from nest sites. Males may sometimes have multiple mates, and bigamy is mostly limited by aggression between females. Many birds do not find a nest and a mate, and instead may serve as helpers around the nest for mated pairs, a role which increases the chances of being chosen to replace a lost mate. Lost mates of both sexes can be replaced quickly during the breeding season. The formation of a pair and the bond between the two birds is tied to the holding of a nest site, though paired house sparrows can recognise each other away from the nest.
House sparrows in natural small populations, as can occur on islands, exhibit inbreeding depression. Inbreeding depression is manifested as lower survival probability and production of fewer offspring, and can occur as a result of the expression of deleterious recessive . In such populations sparrows do not appear to avoid inbreeding.
Especially in warmer areas, the house sparrow may build its nests in the open, on the branches of trees, especially evergreens and hawthorns, or in the nests of large birds such as or . In open nesting sites, breeding success tends to be lower, since breeding begins late and the nest can easily be destroyed or damaged by storms. Less common nesting sites include and , favoured for their warmth; and the old open-topped nests of other songbirds, which are then domed over.
Usually the couples repeat copulation many times. Every copulation is followed by some break of 3 to 4 seconds, and in that time both pair change their position by some distance.
The nest is usually domed, though it may lack a roof in enclosed sites. It has an outer layer of stems and roots, a middle layer of dead grass and leaves, and a lining of feathers, as well as of paper and other soft materials. Nests typically have external dimensions of 20 × 30 cm (8 × 12 in), but their size varies greatly. The building of the nest is initiated by the unmated male while displaying to females. The female assists in building, but is less active than the male. Some nest building occurs throughout the year, especially after moult in autumn. In colder areas house sparrows build specially created roost nests, or roost in street lights, to avoid losing heat during the winter. House sparrows do not hold territories, but they defend their nests aggressively against intruders of the same sex.
House sparrows' nests support a wide range of scavenging insects, including nest flies such as Neottiophilum praestum, Protocalliphora blowflies, and over 1,400 species of beetle.
Some intraspecific brood parasite occurs, and instances of unusually large numbers of eggs in a nest may be the result of females laying eggs in the nests of their neighbours. Such foreign eggs are sometimes recognised and ejected by females. The house sparrow is a victim of interspecific brood parasites, but only rarely, since it usually uses nests in holes too small for parasites to enter, and it feeds its young foods unsuitable for young parasites. In turn, the house sparrow has once been recorded as a brood parasite of the American cliff swallow.
The eggs are white, bluish white, or greenish white, spotted with brown or grey. Subelliptical in shape, they range from in length and in width, have an average mass of , and an average surface area of . Eggs from the tropical subspecies are distinctly smaller. Eggs begin to develop with the deposition of Egg yolk in the ovary a few days before ovulation. In the day between ovulation and laying, egg white forms, followed by eggshell. Eggs laid later in a clutch are larger, as are those laid by larger females, and egg size is hereditary. Eggs decrease slightly in size from laying to hatching. The yolk comprises 25% of the egg, the egg white 68%, and the shell 7%. Eggs are watery, being 79% liquid, and otherwise mostly protein.
The female develops a brood patch of bare skin and plays the main part in avian incubation the eggs. The male helps, but can only cover the eggs rather than truly incubate them. The female spends the night incubating during this period, while the male roosts near the nest. Eggs hatch at the same time, after a short incubation period lasting 11–14 days, and exceptionally for as many as 17 or as few as 9. The length of the incubation period decreases as ambient temperature increases later in the breeding season.
Young house sparrows remain in the nest for 11 to 23 days, normally 14 to 16 days. During this time, they are fed by both parents. As newly hatched house sparrows do not have sufficient insulation, they are brooded for a few days, or longer in cold conditions. The parents swallow the droppings produced by the hatchlings during the first few days; later, the droppings are moved up to away from the nest.
The chicks' eyes open after about 4 days and, at an age of about 8 days, the young birds get their first down feather. If both parents perish, the ensuing intensive begging sounds of the young often attract replacement parents which feed them until they can sustain themselves. All the young in the nest leave it during the same period of a few hours. At this stage, they are normally able to fly. They start feeding themselves partly after 1 or 2 days, and sustain themselves completely after 7 to 10 days, 14 at the latest.
The house sparrow is infested by a number of external parasites, which usually cause little harm to adult sparrows. In Europe, the most common mite found on sparrows is Proctophyllodes, the most common are Argas reflexus and Ixodes arboricola, and the most common flea on the house sparrow is Ceratophyllus gallinae. Dermanyssus blood-feeding mites are also common ectoparasites of house sparrows, and these mites can enter human habitation and bite humans, causing a condition known as gamasoidosis. A number of chewing louse occupy different Ecological niche on the house sparrow's body. Menacanthus lice occur across the house sparrow's body, where they feed on blood and feathers, while Brueelia lice feed on feathers and Philopterus fringillae occurs on the head.
The house sparrow has long been used as a food item. From around 1560 to at least the 19th century in northern Europe, earthenware "sparrow pots" were hung from eaves to attract nesting birds so the young could be readily harvested. Wild birds were trapped in nets in large numbers, and sparrow pie was a traditional dish, thought, because of the association of sparrows with lechery, to have aphrodisiac properties. A traditional Indian medicine, Ciṭṭukkuruvi Lehyam in Tamil language, was sold with similar aphrodisiac claims. Sparrows were also trapped as food for falconry birds and zoo animals. During the 1870s, there were debates on the damaging effects of sparrows in the House of Commons in England. In the early part of the 20th century, sparrow clubs culled many millions of birds and eggs in an attempt to control numbers of this perceived pest, but with only a localised impact on numbers. Sparrows were also persecuted in Germany from at least 1650 until 1970. House sparrows have been kept as pets at many times in history, though they have no bright plumage or attractive songs, and raising them is difficult. The house sparrow has an extremely large range and population, so it is assessed as least concern for conservation on the IUCN Red List.
In Europe, citizen science data from a 21-year study found its numbers had dwindled in its native range in Europe by nearly 60% by 2021. In Great Britain, populations peaked in the early 1970s, but have since declined by 68% overall, and about 90% in some regions. The RSPB lists the house sparrow's UK conservation status as red. In London, the house sparrow almost disappeared from the central city. The numbers of house sparrows in the Netherlands have dropped in half since the 1980s, so the house sparrow is even considered an endangered species. This status came to widespread attention after a female house sparrow, referred to as the "dominomus", was killed after knocking down dominoes arranged as part of an attempt to set a world record. These declines are not unprecedented, as similar reductions in population occurred when the internal combustion engine replaced horses in the 1920s and a major source of food in the form of grain spillage was lost.
Declines have been particularly apparent even in North America, where the house sparrow is invasive in some states. Introduced to Philadelphia initially in 1852 the house sparrow rapidly spread across the nation. However, the bird has largely disappeared from the city nowadays and is estimated to have declined in North America by 84% since 1966.
In South Asia, the house sparrow has largely vanished from many major cities. In general, the house sparrow population has been on the decline in many Asian countries, particularly in India.
Various causes for the dramatic decreases in population have been proposed, including predation, in particular by Eurasian sparrowhawks, possibly facilitated by the elimination of bushes which the sparrows use to hide, electromagnetic radiation from mobile phones; and diseases such as avian malaria. A shortage of nesting sites caused by changes in urban building design is probably a factor, and conservation organisations have encouraged the use of special nest boxes for sparrows. A primary cause of the decline seems to be an insufficient supply of insect food for nestling sparrows. Declines in insect populations result from an increase of monoculture crops, the heavy use of pesticides, the replacement of native plants in cities with introduced plants and parking areas, and possibly the introduction of unleaded petrol, which produces toxic compounds such as methyl nitrite.
Protecting insect habitats on farms and planting native plants in cities benefit the house sparrow, as does establishing urban green spaces. To raise awareness of threats to the house sparrow, World Sparrow Day has been celebrated on 20 March since 2010.
To promote conservation, in 2012, the house sparrow was declared as the state bird of Delhi.
Distribution and habitat
Introduction
Habitat
Behaviour
Social behaviour
Sleep and roosting
Body maintenance
Feeding
Locomotion
Dispersal and migration
Breeding
Nesting
Eggs and young
Survival
Predation
Parasites and disease
Physiology
Relationships with humans
Population decline
Cultural associations
See also
Works cited
External links
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